It is inexplicable on
the theory of creation why a part developed in a very unusual manner in one
species alone of a genus, and therefore, as we may naturally infer, of
great importance to that species, should be eminently liable to variation;
but, on our view, this part has undergone, since the several species
branched off from a common progenitor, an unusual amount of variability and
modification, and therefore we might expect the part generally to be still
variable. But a part may be developed in the most unusual manner, like the
wing of a bat, and yet not be more variable than any other structure, if
the part be common to many subordinate forms, that is, if it has been
inherited for a very long period; for in this case it will have been
rendered constant by long-continued natural selection.
Glancing at instincts, marvellous as some are, they offer no greater
difficulty than do corporeal structures on the theory of the natural
selection of successive, slight, but profitable modifications. We can thus
understand why nature moves by graduated steps in endowing different
animals of the same class with their several instincts. I have attempted
to show how much light the principle of gradation throws on the admirable
architectural powers of the hive-bee. Habit no doubt often comes into play
in modifying instincts; but it certainly is not indispensable, as we see in
the case of neuter insects, which leave no progeny to inherit the effects
of long-continued habit.
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