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Darwin, Charles, 1809-1882

"The Origin of Species by means of Natural Selection, 6th Edition"


Considering the several rules now given, which govern the fertility of
first crosses and of hybrids, we see that when forms, which must be
considered as good and distinct species, are united, their fertility
graduates from zero to perfect fertility, or even to fertility under
certain conditions in excess; that their fertility, besides being eminently
susceptible to favourable and unfavourable conditions, is innately
variable; that it is by no means always the same in degree in the first
cross and in the hybrids produced from this cross; that the fertility of
hybrids is not related to the degree in which they resemble in external
appearance either parent; and lastly, that the facility of making a first
cross between any two species is not always governed by their systematic
affinity or degree of resemblance to each other. This latter statement is
clearly proved by the difference in the result of reciprocal crosses
between the same two species, for, according as the one species or the
other is used as the father or the mother, there is generally some
difference, and occasionally the widest possible difference, in the
facility of effecting an union. The hybrids, moreover, produced from
reciprocal crosses often differ in fertility.
Now do these complex and singular rules indicate that species have been
endowed with sterility simply to prevent their becoming confounded in
nature? I think not.


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