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Darwin, Charles, 1809-1882

"The Origin of Species by means of Natural Selection, 6th Edition"

It must not be inferred from these remarks that any of the
grades of wing-structure here alluded to, which perhaps may all be the
result of disuse, indicate the steps by which birds actually acquired their
perfect power of flight; but they serve to show what diversified means of
transition are at least possible.
Seeing that a few members of such water-breathing classes as the Crustacea
and Mollusca are adapted to live on the land; and seeing that we have
flying birds and mammals, flying insects of the most diversified types, and
formerly had flying reptiles, it is conceivable that flying-fish, which now
glide far through the air, slightly rising and turning by the aid of their
fluttering fins, might have been modified into perfectly winged animals.
If this had been effected, who would have ever imagined that in an early
transitional state they had been inhabitants of the open ocean, and had
used their incipient organs of flight exclusively, so far as we know, to
escape being devoured by other fish?
When we see any structure highly perfected for any particular habit, as the
wings of a bird for flight, we should bear in mind that animals displaying
early transitional grades of the structure will seldom have survived to the
present day, for they will have been supplanted by their successors, which
were gradually rendered more perfect through natural selection.
Furthermore, we may conclude that transitional states between structures
fitted for very different habits of life will rarely have been developed at
an early period in great numbers and under many subordinate forms.


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